Females cats will sometimes do it when they are in heat, but generally it is mostly un-neutured male tom cats.
Cats spray for a variety of reasons. They spray during territorial disputes, when they are aware of another cat in heat, when stressed and often after fights with other cats. So the cats, normally solitary creatures, mark their territory as a way to reduce conflicts with other cats in the area. Mostly cats will spray outside, but if there are conflicts
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